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In parallel, increased activity of lipid metabolic enzymes, including acyl-CoA synthetase long-chain family member 4 ( ACSL4 ) and lysophosphatidylcholine acyltransferase 3 ( LPCAT3 ), promotes the enrichment and remodeling of polyunsaturated fatty acids (PUFAs) in membrane phospholipids (Yang and Stockwell, 2016